In botanya berry is a fleshy fruit without a stone produced from a single flower containing one ovary. Berries so defined include grapescurrantsand tomatoesas well as cucumberseggplants aubergines and bananasbut exclude certain fruits commonly called berries, such as strawberries and raspberries. The berry is the most common type of fleshy fruit in
De fructibus et seminibus plantarum online dating the De fructibus et seminibus plantarum online dating outer layer of the ovary wall ripens into a potentially edible " pericarp ".
Berries may be formed from one or more carpels from the same flower i. The seeds are usually embedded in the fleshy interior of the ovary, but there are some non-fleshy exceptions, such as pepperswith air rather than pulp around their seeds. Many berries are edible, but others, such as the fruits of the potato and the deadly nightshadeare poisonous to humans.
Some berries, such as the white and red mulberryare poisonous when unripe, but are edible in their ripe form. A plant that bears berries is said to be bacciferous or baccate a fruit that resembles a berry, whether it actually is a berry or not, can also be called "baccate". In everyday English, a " berry " is any small edible fruit. Berries are usually juicy, round, brightly coloured, sweet or sourand do not have a stone or pit, although many pips or seeds may be present. In botanical language, a berry is a simple fruit having seeds and fleshy pulp the pericarp produced from the ovary of a single flower.
The ovary can be inferior or superior. It is indehiscenti. The outer layer is called the "exocarp" or " epicarp "; the middle layer, the " mesocarp " or "sarcocarp"; the inner layer, the " endocarp ".
Botanists have not applied these terms consistently. Exocarp and endocarp may be restricted to more-or-less single-layered "skins", or may include tissues adjacent to them; thus on one view, the exocarp extends inwards to the layer of vascular bundles "veins".
The inconsistency in usage has been described as "a source
De fructibus et seminibus plantarum online dating confusion". The nature of the endocarp distinguishes a berry from a drupe, which has a hardened or stony endocarp see also below.
The two kinds of fruit intergrade, depending on the state of the endocarp. Some sources have attempted to quantify the difference, e. This distinguishes, for example, a Vaccinium or Solanum berry from an Adansonia baobab amphisarca, which has a dry, more rigid and self-supporting skin.
Berries that develop from an inferior ovary are sometimes termed epigynous berries or false berries, as opposed to true berries, which develop from a superior ovary. In epigynous De fructibus et seminibus plantarum online dating, the berry includes tissue derived from parts of the flower besides the ovary.
The floral tube, formed from the basal part of the sepals, petals and stamens can become fleshy at maturity and is united with the ovary to form the fruit. Common fruits that are sometimes classified as epigynous berries include bananascoffeemembers of the genus Vaccinium e. Many fruits commonly referred to as berries are not actual berries by the scientific definition, but fall into one of the following categories:.
Drupes are fleshy fruits produced from a usually single-seeded ovary with a hard woody layer called the endocarp surrounding the seed. Familiar examples include the stonefruits of the genus Prunus peachesplums and cherriesolivescoconutbayberry and Persea species.
Some definitions make the mere presence of an internally differentiated endocarp the defining feature of a drupe;  others qualify the nature of the endocarp required in a drupe, e. Other drupe-like fruits with a single seed that lack the stony endocarp include sea-buckthorn Hippophae rhamnoidesElaeagnaceaewhich is an achenesurrounded by a swollen hypanthium that provides the fleshy layer.
The pome fruits produced by plants in subtribe Pyrinae of family Rosaceaesuch as apples and pears, have a structure the core in which tough tissue clearly separates the seeds from the outer softer pericarp. Amelanchier pomes become so soft at maturity that they resemble a blueberry and are known as Juneberries, serviceberries or Saskatoon berries.
Aggregate or compound fruits contain seeds from different ovaries of a single flower, with the individual "fruitlets" joined together at maturity to form the
De fructibus et seminibus plantarum online dating fruit.
Multiple fruits are the fruits of two or more multiple flowers that are merged or packed closely together. The female seed cones of some conifers have fleshy and merged scales, giving them a berry-like appearance. Juniper "berries" family Cupressaceaein particular those of Juniperus communisare used to flavour gin. The seed cones of species in the Podocarpaceae and Taxaceae families have a bright colour when fully developed, increasing the resemblance to true berries.
The "berries" of yews Taxus species consist of a female seed cone with which develops a fleshy red aril partially enclosing the poisonous seed. The Latin word baca or bacca plural baccae was originally used for "any small round fruit". He did not make the modern distinction between "fruits" and "seeds", calling hard structures like nuts semina or seeds. A fleshy fruit was called a pericarpium.
For Caesalpinus, a true bacca or berry was a pericarpium derived from a flower with a superior ovary; one derived from a flower with an inferior ovary was called a pomum.
InCarl Linnaeus wrote Philosophia Botanicaconsidered to be the first textbook of descriptive systematic botany. Botanists continue to differ on how fruit should be classified.
Joseph Gaertner published a two-volume "De fructibus et seminibus plantarum online dating," De Fructibus et Seminibus Plantarum on the fruits and seeds of plants between and In addition to Linnaeus' eight terms, he introduced seven more, including pepo for the berry-like fruits of cucurbits. Nicaise Auguste Desvaux in used the terms hesperidium and amphisarca as further subdivisions of berries.
An amphisarca was described as woody on the outside and fleshy on the inside. There remains no universally agreed system of classification for fruits, and there continues to be "confusion over classification of fruit types and the definitions given to fruit terms".
By definition, berries have a fleshy, indehiscent pericarp, as opposed to a dry, dehiscent pericarp. Fossils show that early flowering plants had dry fruits; fleshy fruits, such as berries or drupes, appeared only towards the end of the Cretaceous Period or the beginning of the Paleogene Periodabout 66 million years ago. The increasing importance of seed dispersal by fruit-eating vertebrates, both mammals and birds, may have driven the evolution of fleshy fruits.
Alternatively, the causal direction may be the other way round. Large fleshy fruits are associated with moist habitats with closed tree canopies, where wind dispersal of dry fruits is less effective. Such habitats were increasingly common in the Paleogene and the associated change in fruit type may have led to the evolution of fruit eating in mammals and birds. Fruit type has been considered to be a useful character in classification and in understanding the phylogeny of plants.